Why is "nuq" attached to "jatlh"? After quickly checking my copy of Marc Okrand's The Klingon Dictionary, it seems like this should have been "nuq Dajatlh mara?"
You are correct that a formal way of saying, "What did you say, Mara?" would be "nuq Dajatlh, mara?" However, if you look up "nuqjatlh" in the dictionary portion of the addendum (page 183), you will see that it is an exclamation meaning, "what did you say? huh? what?". Since students are still learning the grammar at this point, we have decided to introduce them to this useful exclamation early in the course.
Hmm....is there any significant difference in pronunciation?
And what if you are writing in piqaD, in which spaces between words are optional?